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A drop of sunlight falls to the ground and grows into a magical flower, with the power to heal the sick and injured. An old woman named Gothel finds it and uses it to keep herself young by singing an incantation. Centuries later, the Queen becomes ill during childbirth, and so the flower is found. She is healed and gives birth to a daughter, whom they name Rapunzel, whose golden hair is now the source of the magical power. Gothel kidnaps Rapunzel and hides her in a tall tower, raising her as her own child. Gothel knows that if Rapunzel's hair is cut, it will turn brown and lose its magic, so Rapunzel's hair is left to grow. Every year, on Rapunzel's birthday, her kingdom releases thousands of sky lanterns, in the hope that the lost princess will return. On the eve of her eighteenth birthday, Rapunzel asks Gothel to let her see the floating lights, but Gothel refuses.
Meanwhile, a group of thieves, led by the charismatic Flynn Rider, steals the tiara of the lost princess from the castle. During the ensuing chase, a horse named Maximus is separated from his rider, the Captain of the Guards, and continues the search for Flynn on his own. Flynn outwits his accomplices and takes the tiara; he abandons them and stumbles upon Rapunzel's tower. Once inside the tower, Rapunzel knocks Flynn unconscious with a frying pan. She hides Flynn inside her closet and confiscates the tiara. When Gothel returns, Rapunzel requests, as a birthday present, a special paint made from white shells from a faraway beach. Gothel leaves on the three-day journey to bring back the shells. Rapunzel makes a deal with Flynn: a journey to the "lights" in exchange for the tiara. Flynn attempts to make Rapunzel end their journey by taking her to the Snuggly Duckling inn, which is full of Gaul thugs, but the thugs are charmed by Rapunzel, who encourages them to follow their dreams.
Mother Gothel sees Maximus riderless and worries someone will find Rapunzel. She returns to the tower to find Rapunzel gone. Meanwhile, the guards invade the tavern, but Rapunzel and Flynn have escaped. The pursuit ends at a dam, which Maximus causes to collapse; Flynn and Rapunzel are trapped in a flooding cave. Believing he's about to die, Flynn admits his true name: Eugene Fitzherbert. Rapunzel admits she has magic hair that glows when she sings. Using her power, they find a way out. Later, when Flynn goes to gather firewood, Gothel secretly meets Rapunzel. Gothel tells Rapunzel that Flynn doesn't care for her and merely wants the tiara. Gothel gives Rapunzel the tiara, insisting that she tests Flynn by giving it to him.
The next morning, Maximus confronts Flynn, but to Flynn's dismay Rapunzel befriends the horse. The three arrive at the kingdom and that night Flynn takes Rapunzel to see the lanterns. There, Rapunzel gives Flynn back the tiara. Flynn spies the Stabbington brothers and leaves Rapunzel waiting as he gives them the tiara back. However, the brothers tie him up on a boat and sail him across the lake. They reveal Flynn's "betrayal" to Rapunzel as they attempt to kidnap her for her hair's power, but Gothel "rescues" her and they return to the tower. Later, Flynn is arrested and sentenced to death. Maximus brings the inn thugs to rescue Flynn, and Maximus and Flynn race back to the tower. From various clues she found during her adventure, Rapunzel realizes she is the lost princess and attempts to flee the tower. When Flynn escapes on Maximus and returns to the tower, he climbs up Rapunzel's hair only to find her chained to the wall and gagged. Gothel stabs him from behind and prepares to take Rapunzel to a new hiding place, detatching the end of the chain from the wall and trying to pull her away. But Pascal, Rapunzel's pet Chameleon, struggles with Gothel. Rapunzel's gag comes off and she says she will always fight, but promises to go with Gothel willingly if she allows her to heal Flynn. Knowing that Rapunzel keeps her word, Gothel agrees, but before Rapunzel heals him, Flynn cuts her hair with a shard of broken mirror. Rapunzel's hair turns brown and loses its power and Gothel rapidly begins to age. Disoriented, Gothel stumbles around the tower when Pascal causes her to trip and fall from the tower. She turns to dust before she hits the ground.
Flynn slowly dies in Rapunzel's arms. Heartbroken, she cries, and tearfully sings the incantation. One teardrop, filled with her power, lands on his cheek and revives him. The two embrace and kiss. Back at the kingdom, the royal family has a tearful reunion. Years later, Flynn and Rapunzel are married. Along the way, the Gaul thugs fulfill their individual dreams, and Maximus becomes a respected official on the Royal Guard.
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